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Environmental factors, genetic predisposition, and smoking are significant contributors to the development of RA ( In recent decades, the incidence of RA has increased significantly, and this trend is expected to continue ( This chronic condition is characterized by inflammation of the synovial membrane, leading to joint damage, swelling, stiffness, pain, tissue degradation, deformity, and instability

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Supplementary information Supplementary Table 1 (download XLSX Proteomics, cryo-EM data collection, genetic screens and RNA-Seq